Many factors. Foremost: water chemistry imbalance that leads to sustained stress and its underlying cause is typically due to a filter needing major attention. or undersized. Foul brine-shrimp is the foremost factor in fact if you give them brine shrimp which has a stench to it, you can almost guarantee that within the week, you'll see increasing numbers of lifeless bodies beginning to litter the bottom (unless the parents get to it first). This last one is the most common cause of die-offs in fish under dime size. Unfortunately, once they begin dropping like flies, those already affected will not respond to therapies/interventions. If anything, medications become highly toxic for them to handle and accelerate their demise. However, interventions will save those yet unaffected ones.
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